Glossary of "Evolution (4th ed.)" by Futuyma & Kirkpatrick (2017)
- Any reduction of the mean fitness of a population resulting from the existence of genotypes with a fitness lower than that of the most fit genotype.
- Futuyma, D. J. & Kirkpatrick, M. 2017. Evolution, 4th ed. -xviii + 602 pp. Sinauer Associates, Inc., Sunderland.