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== Definition ==
== Definition ==
=== Glossary of Plant Systematics (1st ed.) by [[Simpson_2006|Simpson (2006)]] ===
* The outer, protruding layers of the exine, including columellae, tectum, and supratectal sculpturing elements.
=== Glossary of "The Evolution of Plants (2nd ed.)" by [[Willis_McElwain_2014|Willis & McElwain (2014)]] ===
=== Glossary of "The Evolution of Plants (2nd ed.)" by [[Willis_McElwain_2014|Willis & McElwain (2014)]] ===
* The outer layer of the [[exine]] in pollen grains.
* The outer layer of the [[exine]] in pollen grains.