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*Hara, M. & Miyakuni, Y. 1991. Hepaticae and Anthocerotae of Tanegashima Island, Kyusyu, Japan. Mem. Fac. Sci. Kochi Univ. Ser. D. Biol. 12: 1–10.
 
*Hara, M. & Miyakuni, Y. 1991. Hepaticae and Anthocerotae of Tanegashima Island, Kyusyu, Japan. Mem. Fac. Sci. Kochi Univ. Ser. D. Biol. 12: 1–10.
 
*Miyagi, Y. 1977. On the limestone flora of Kikai Island, the Amami Islands. Ecological studies of nature conservation of Ryukyu Islands (III): 23–28.
 
*Miyagi, Y. 1977. On the limestone flora of Kikai Island, the Amami Islands. Ecological studies of nature conservation of Ryukyu Islands (III): 23–28.
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*宮本光生. 1960. 屋久島の蘚類. 屋久島の生物調査報告: 23–26.
 
*Yokoyama, H., Yamaguchi. T., Nishimura, N., Furuki, T. & Akiyama, H. 2007. Checklist of bryophytes known from Yakushima Island, southern Kyushu, Japan. Bryol. Res. 9: 159-197.
 
*Yokoyama, H., Yamaguchi. T., Nishimura, N., Furuki, T. & Akiyama, H. 2007. Checklist of bryophytes known from Yakushima Island, southern Kyushu, Japan. Bryol. Res. 9: 159-197.
  
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