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== Definition ==
== Definition ==
=== Glossary of "Evolution (4th ed.)" by [[Futuyma_Kirkpatrick_2017|Futuyma & Kirkpatrick (2017)]] ===
=== Glossary of "Evolution (4th ed.)" by [[Futuyma_Kirkpatrick_2017|Futuyma & Kirkpatrick (2017)]] ===
*The existence of both a fitness benefit and a fitness cost of a mutation or character state, relative to another.
*The existence of both a fitness benefit and a fitness cost of a [[mutation]] or [[character_state|character state]], relative to another.
==文献（引用）==
==文献（引用）==
