Glossary of Plant Systematics (1st ed.) by Simpson (2006)
- A polymer of fatty acids, functioning as a sealant in the cuticle layer of land plants, inhibiting water loss.
Glossary of "Evolution of Plants (2nd ed.)" by Willis & McElwain (2014)
- An important chemical component of plant cuticle, which is less resistant to decay than cutan and is therefore not detected as often as cutan in the fossil record.
- Simpson, M. G. 2006. Plant Systematics. -xi + 590 pp. Elsevier Academic Press, Oxford.
- Willis, K. J. & McElwain, J. C. 2014. The Evolution of Plants, 2nd ed. -ix + 398 pp. Oxford University Press, Oxford.