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Definition
Glossary of "Plant Biology (1st ed.)" by Graham et al. (2003)
- Flowers that possess all four floral whorls: sepals, petals, stamens, and carpels.
Glossary of Plant Systematics (1st ed.) by Simpson (2006)
- Having all four major whorls or floral parts: sepals, petals, stamens, carpels. (flower cycly)
文献（引用）
- Graham, L. E., Graham, J. M. & Wilcox, L. W. 2003. Plant Biology. -xxxv + 497 pp. Pearson Education, Inc., Upper Saddle River.
- Simpson, M. G. 2006. Plant Systematics. -xi + 590 pp. Elsevier Academic Press, Oxford.
